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Cited article:

Homeosis and delayed floral meristem termination could account for abnormal flowers in cultivars of Delphinium and Aquilegia (Ranunculaceae)

Felipe Espinosa, Catherine Damerval, Martine Le Guilloux, Thierry Deroin, Wei Wang, Myreya Pinedo-Castro, Sophie Nadot and Florian Jabbour
Botanical Journal of the Linnean Society 195 (3) 485 (2021)
https://doi.org/10.1093/botlinnean/boaa063